Biomolecules β the chemistry of life
Every living cell is built from four classes of macromolecules, all polymers of smaller monomers.
| Biomolecule | Monomer | Key bond | Main function |
|---|---|---|---|
| Carbohydrate | Monosaccharide | Glycosidic | Energy, structure (cellulose) |
| Protein | Amino acid | Peptide | Enzymes, structure, transport |
| Lipid | Glycerol + fatty acid | Ester | Membranes, energy store |
| Nucleic acid | Nucleotide | Phosphodiester | Information storage (DNA/RNA) |
- Carbohydrates: general formula (CHβO)β. Glucose (CβHββOβ) is the primary respiratory substrate. Cellulose (Ξ²-1,4 glucose) is the most abundant organic polymer on Earth and forms the plant cell wall; starch (Ξ±-glucose) is the storage form.
- Proteins: 20 standard amino acids joined by peptide bonds. Four levels of structure β primary, secondary (Ξ±-helix, Ξ²-pleated sheet), tertiary, quaternary. Enzymes are globular proteins that lower activation energy.
- Lipids: glycerol esterified to fatty acids; phospholipids form the amphipathic bilayer of membranes.
- Nucleic acids: DNA is double-stranded (deoxyribose, A-T-G-C); RNA is usually single-stranded (ribose, A-U-G-C).
High-yield: Water is the universal solvent and the medium of all metabolism; its high specific heat, cohesion, and surface tension drive processes like the transpiration pull.
The cell as the unit of life
The cell theory (Schleiden, Schwann, Virchow) states: all organisms are made of cells, the cell is the structural and functional unit of life, and all cells arise from pre-existing cells (Omnis cellula-e-cellula). Prokaryotes (bacteria, cyanobacteria) lack a true nucleus and membrane-bound organelles; eukaryotes possess both.
Five-kingdom classification
R. H. Whittaker (1969) proposed five kingdoms based on cell type, body organisation, mode of nutrition and phylogeny.
| Kingdom | Cell type | Nutrition | Example |
|---|---|---|---|
| Monera | Prokaryotic | Auto/heterotrophic | Bacteria, cyanobacteria |
| Protista | Eukaryotic (unicellular) | Photo/holozoic | Chlamydomonas, diatoms |
| Fungi | Eukaryotic | Saprophytic/parasitic | Mushroom, yeast |
| Plantae | Eukaryotic | Autotrophic | Mosses, ferns, trees |
| Animalia | Eukaryotic | Holozoic (heterotrophic) | All animals |
- Two-kingdom system (Linnaeus) split life into Plantae and Animalia only; it failed to separate prokaryotes from eukaryotes.
- Binomial nomenclature (Linnaeus): every species gets a two-word Latin name β genus (capitalised) + species (lower case), italicised, e.g. Oryza sativa (rice).
Plant diversity β from algae to angiosperms
Kingdom Plantae is arranged in ascending complexity:
- Algae (Thallophyta): simple, autotrophic, mostly aquatic; no true roots/stems/leaves. Grouped by pigment β green (Chlorophyceae), brown (Phaeophyceae), red (Rhodophyceae). Reserve food and cell-wall material differ per group.
- Bryophyta (mosses, liverworts): "amphibians of the plant kingdom". Dominant gametophyte (haploid); lack vascular tissue; need water for fertilisation. Example: Funaria, Marchantia.
- Pteridophyta (ferns): first true vascular plants with xylem and phloem. Dominant sporophyte (diploid); reproduce by spores. Example: Dryopteris, Selaginella.
- Gymnosperms: naked seeds (no ovary/fruit); mostly evergreen; well-developed vascular tissue. Example: Pinus, Cycas, Ginkgo.
- Angiosperms: flowering plants with seeds enclosed in fruit; show double fertilisation and vascular tissue with vessels. Divided into dicots and monocots.
Alternation of generations
Land plants alternate between a haploid gametophyte (produces gametes by mitosis) and a diploid sporophyte (produces spores by meiosis). The trend across evolution is progressive reduction of the gametophyte and dominance of the sporophyte.
Dicots vs monocots
| Feature | Dicot | Monocot |
|---|---|---|
| Cotyledons | Two | One |
| Leaf venation | Reticulate | Parallel |
| Root | Tap root | Fibrous |
| Floral parts | Tetra/pentamerous | Trimerous |
| Vascular bundles | Ring, open | Scattered, closed |
Biodiversity concepts
- Biodiversity = variety of life at three levels: genetic, species, and ecosystem diversity.
- Nepal, despite its small area, holds exceptional biodiversity because of its steep altitudinal gradient (60 m to 8,848 m), spanning tropical to alpine zones.
- Hotspots are regions rich in endemic species yet under threat; the Himalaya is a recognised global biodiversity hotspot.
- Conservation is in-situ (national parks, biosphere reserves, sacred groves) or ex-situ (botanical gardens, seed banks, tissue culture).
Exam tip: Remember the diagnostic "firsts" β bryophytes = first land plants, pteridophytes = first vascular plants, gymnosperms = first seed plants, angiosperms = first flowering plants with double fertilisation.
Quick revision
- Cellulose (Ξ²-1,4 glucose) is the most abundant organic polymer and forms the plant cell wall.
- Whittaker's five kingdoms (1969): Monera, Protista, Fungi, Plantae, Animalia.
- Binomial nomenclature by Linnaeus β genus + species, italicised.
- Bryophytes are "amphibians of the plant kingdom"; gametophyte dominant.
- Pteridophytes = first vascular plants; gymnosperms bear naked seeds.
- Angiosperms show double fertilisation and enclosed seeds (fruit).
- Dicot: reticulate venation, tap root, tetra/pentamerous flowers; monocot is the reverse.
- Biodiversity operates at genetic, species and ecosystem levels.